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The conserved catalytic site comprises a tetrad of amino acids: Sec46, Gln81, Trp136 and Asn147, the latter being directly involved in catalysis

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These findings indicate that repeated agonist treatment initiates a certain signaling pathway that promotes the USP33-mediated deubiquitination of arrestin3, and that this deubiquitinated arrestin3 subsequently undergoes complexation with G, which in turn translocates to the nucleus via an importin complex (Fig